Mass Effect is set within the Milky Way galaxy in the year 2183 where interstellar travel is possible through the use of mass transit devices called Mass Relays, a technology believed to have been built by an extinct alien race known as the Protheans. The term "Mass Effect" is defined as a form of mass-negating technology, allowing the creation of physics phenomena, like artificial gravity or FTL travel. A conglomerate body of governments known as the Citadel Council controls a large percentage of the galaxy and is responsible for maintaining law and order among races of the galactic community. Races that belong to the Citadel Council include humans, asari, salarians, and turians. Other alien races seen in the game include the reptilian krogan, the environmental suited quarians, and a hostile race of networked artificial intelligences called geth. Humanity is represented by the Systems Alliance, an organized body that joined the Citadel Council in 2157.
Shepard and Kaidan are sent to the human colony of Eden Prime to recover an unearthed Prothean beacon. They repel an attack by the geth, rescuing Ashley in the process. Rogue turian Spectre, Saren Arterius, activates the beacon before escaping. Shepard reaches the beacon and receives a vision showing scenes of war and death. The SSV Normandy and its crew are summoned by Ambassador Donnel Udina to the Citadel Station, but Shepard is unable to convince the Citadel Council of Saren's treason without solid evidence. Shepard meets Garrus and Wrex, and rescues Tali, who possesses a recording of a conversation between Saren and asari Matriarch, Benezia. The two discuss their victory while also mentioning an artifact called the "Conduit" and the return of the Reapers, a highly advanced machine race of synthetic-organic starships believed to eradicate all organic civilization every 50,000 years. The Council revokes Saren's Spectre status and makes Shepard the first human Spectre. Shepard is also handed command of the Normandy by Anderson, and begins to follow several leads provided by him and Udina.
On the world of Therum, Shepard rescues Benezia's daughter Liara T'Soni, who joins Shepard's squad because of her biotic abilities and expertise of the Protheans. On the colony of Feros, Shepard fights off Saren's forces and learns that his flagship, Sovereign, possesses unique mind-control capabilities. On the world of Noveria, Shepard tracks down and defeats Benezia, who reveals that she and Saren are being indoctrinated by Sovereign. After completing these missions, the Council informs Shepard that a salarian infiltration unit has uncovered Saren's main base on Virmire. Upon arrival, Shepard learns that Saren has discovered a cure for the krogan genetic disease, the genophage, and plans to breed an army of unstoppable krogan warriors. Wrex clashes with Shepard over whether to destroy it, which results in him either standing down or being executed. After this conflict, Shepard assists the salarians in destroying the base by planting a bomb in it. Inside, Shepard is confronted by Sovereign, who reveals itself to be an actual Reaper.
Sovereign reveals that the Reapers remain outside the Milky Way waiting for organic life to develop and discover the Mass Relays, before harvesting them when they reach their peak. Afterwards, with the bomb about to explode, Shepard hears that Ashley and Kaidan have both been pinned down, and must choose which one to rescue. Shepard also finds Saren, who claims that his allegiance to Sovereign will save organic life forms by demonstrating their usefulness to the Reapers. Liara is then able to pinpoint the Conduit's location back on the Normandy: a Prothean world known as Ilos. There, Shepard learns that the Citadel Station is actually a huge Mass Relay that the Reapers use to invade the galaxy. During the last extinction cycle, a few Protheans survived on Ilos via cryopreservation and then re-entered the Citadel Station via the Conduit, a reverse-engineered miniature Mass Relay disguised aboard the station as a statue. The Protheans sabotaged the Citadel to prevent the Reapers from remotely activating the Citadel Mass Relay. Saren plans to reach the Citadel by using the Conduit to allow Sovereign to activate the Citadel Mass Relay and trigger the Reaper invasion.
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